David Guttenfelder was born in Iowa and graduated from the University of Iowa with a Bachelor of Arts in cultural anthropology. He has worked for the AP since 1994 based in Kenya, the Ivory Coast, India and Japan. He is currently AP’s chief Asia photographer and his feature work has been used prominently in editorial publications throughout the world including Time and National Geographic. Photo: U.S. soldiers from the 2nd Battalion, 12th Infantry Regiment, 4th Brigade Combat Team, fire mortars at known enemy firing positions from a base in the Pech River Valley in Afghanistan's Kunar province, Saturday, October 24, 2009. (Photo by David Guttenfelder/AP Photo)

Polar frame, by Dmitry Kokh, Russia. When Kokh’s boat approached the small island of Kolyuchin in the Russian High Arctic, which had been abandoned by humans since 1992, he was surprised to spot movement in one of the houses. Binoculars revealed polar bears – more than 20 in total – exploring the ghost town. Dmitry used a low-noise drone to document them. (Photo by Dmitry Kokh/Wildlife Photographer of the Year 2022)

Shortlisted: The Life: A narrow strip, Río Tinto, Huelva, Spain. A narrow strip of road divides ochre-coloured and fresh waters. Trees wait on the fresh side of the road opposite the toxic waters of a nearby mine reservoir. (Photo by Roberto Bueno/CIWEM Environmental Photographer of the Year 2021)

Guardian of the Mangroves – Overall Winner. Tanya Houppermans, Cuba. A curious American crocodile (Crocodylus acutus) swims right up to Tanya, at Gardens of the Queen (Jardines De La Reina), an archipelago off the coast of Cuba. It has been strictly protected since 1996, and is one of the most untouched marine ecosystems in the world. ‍ “The healthy population of American crocodiles is down to the pristine condition of the mangroves and I wanted to capture close ups of this gentle giant in its natural habitat. I hope this image can illustrate that protecting areas like this is so critical”. (Photo by Tanya Griffin Houppermans/Mangrove Photographer of the Year)

Youth category winner: Symphony No 5 by Giacomo Redaelli (Italy). (Photo by Giacomo Redaelli/2019 Nature Photographer of the Year)

This photo was caught on camera by amateur photographer Kutub Uddin, 27, and was taken in his back garden in Bognor Regis, West Sussex. The red-eyed tree frogs have been Mr Uddin's pets for four months and he often lets them out to roam around his garden. Photo: “Ohh no :( Raining”. (Photo by Kutub Uddin)

The CBRE Urban Photographer of the Year competition – now in its ninth year – encourages professional and amateur photographers to capture cities at work. Here: Dog Walker – 15:00. The competition allows photographers to enter up to 24 images, one representing each hour of the day. (Photo by Johanna Siegmann)

Respect, Kronotsky nature reserve, Russia. The photographer’s cat, Ryska – her name means little lynx in Russian – stands outside their cabin and with aggressive posturing warns off a fox. In winter, foxes would regularly visit the cabin searching for food. If one peered in at the window, possible when the snow was deep, Ryska would sit on the other side, fur raised, and growl. When outside, she would hold her ground. The foxes were not always frightened and so encounters could be a sort of dance. (Photo by Igor Shpilenok/Unforgettable Behaviour/NHM)