Shooting star by Tony Wu, US/Japan. Winner, underwater. The electrifying reproductive dance of a giant sea star. As the surrounding water filled with sperm and eggs from spawning sea stars, Wu faced several challenges. Stuck in a small, enclosed bay with only a macro lens for photographing small subjects, he backed up to squeeze the undulating sea star into his field of view. The dancing posture of spawning sea stars rising and swaying may help release eggs and sperm, or may help sweep the eggs and sperm into the currents where they fertilise together in the water. Kinko Bay, Japan. (Photo by Tony Wu/Wildlife Photographer of the Year)

A huge atlas moth, with a wingspan of more than 9in, photographed on an areca nut plantation in Sirsi, India. (Photo by Uday Hegde/Close Up Photographer of the Year)

Performers eat their lunch before the start of the New Year's Day Parade in London, January 1, 2016. (Photo by Kevin Coombs/Reuters)

Michael O’Neill won a prize in animal portraits with fry of a peacock bass hovering around their mother for protection against predators in South Florida. (Photo by Michael Patrick O'Neill/2016 National Geographic Nature Photographer of the Year)

A Ukrainian serviceman fires a 2S7 Pion self-propelled gun during military exercises near the village of Divychky in Kiev region, Ukraine, October 21, 2016. (Photo by Valentyn Ogirenko/Reuters)

Model Kendall Jenner poses on the red carpet as she arrives for the screening of the film “Mal de pierres” (From the Land of the Moon) in competition at the 69th Cannes Film Festival in Cannes, France, May 15, 2016. (Photo by Yves Herman/Reuters)

A dirty thunderstorm occurs over Mount Sakurajima as the volcano erupts violently at 12:03 am on July 26, 2016 in Tarumizu, Kagoshima, Japan. The eruption occurred at the mountain's Showa crater. This is the first time that the active volcano in southern Kyushu has spewed out a smokestack that high since an eruption on August 18, 2013, according to the Kagoshima Meteorological Office. (Photo by The Asahi Shimbun via Getty Images)

Fireworks explode over the Kremlin in Red Square which was blocked by police during New Year celebrations in Moscow, Russia, Sunday, January 1, 2017. New Year's Eve is Russia's major gift-giving holiday, and big Russian cities were awash in festive lights and decorations. (Photo by Alexander Zemlianichenko Jr./AP Photo)