Winner – Animal Portraits: The pose by Mogens Trolle, Denmark. A young male proboscis monkey cocks his head slightly and closes his eyes. Unexpected pale blue eyelids now complement his immaculately groomed auburn hair. He poses for a few seconds as if in meditation. He is a wild visitor to the feeding station at Labuk Bay Proboscis Monkey Sanctuary in Sabah, Borneo – “the most laid-back character”, says Trolle, “quite unlike anything I’ve ever seen on another monkey” – connects us, he hopes, with a fellow primate. (Photo by Mogens Trolle/Wildlife Photographer of the Year 2020)

Buddhist monk Wilatha holds a rescued Burmese python at his monastery that has turned into a snake sanctuary on the outskirts of Yangon, Myanmar, November 26, 2020. Wilatha is trying to play a part in saving scores of snakes that might otherwise be killed or destined for the black market. (Photo by Shwe Paw Mya Tin/Reuters)

A butterfly is seen at a farm in Beykoz disrtict of Istanbul, Turkey on October 26, 2021. Cigdem Unlu runs a farm where 800 butterflies live and being presented to the visitors as the environment exclusively weathered for butterflies. (Photo by Elif Ozturk Ozgoncu/Anadolu Agency via Getty Images)

“After an afternoon of hiking and exploring the caves of Rhossili Bay, West Glamorgan, we caught the sun breaking through the clouds”. (Photo by Megan Warren-Davis/The Guardian)

Taking it all in at Trolltunga, a piece of rock which juts horizontally out of a mountain, 700 metres above lake Ringedalsvatnet in Norway. (Photo by Sam Rogers/GuardianWitness)

These incredible pictures look like a bird's-eye view of a faraway alien planet – but they're actually hot springs. The amazing snaps are up-close shots of Iceland's hot springs, and the point-blank pictures produce a breathtaking array of colors and effects. Here: The amazing snaps are up-close shots of Icelandic hot springs. (Photo by Johann Vilhjalmsson/Caters News)

Runner-up. “The heavens opened and the streets around Shibuya, Tokyo, were suddenly drenched in even more colour and movement. I was taking photos (my friend was holding the umbrella) when I noticed this woman at the crossing. I’m pleased with the futuristic feel of the image”. (Photo by Katherine Bridgestock/The Guardian)

Neil Zeller's snaps were taken across Canada, where he photographs the northern lights dancing across a dark night sky. In one stunning shot, Zeller, 42, even manages to capture a flash of lightning beneath a storm cell while an aurora gives off a magnificent green glow. The photographer, from Calgary, Alberta, describes himself as a night person, and became interested in night photography about 10 years ago. (Photo by Neil Zeller/Caters News)