Winner. “I took this vertical image in the Quarry Bay district of Hong Kong during the dusk ‘blue hour’, when there was a perfect balance between the ambient light in the sky and the artificial lights of the high-rise residential buildings”. (Photo by Jatinder Heer/The Guardian)

Behaviour winner; Courting Royals: two royal terns in courtship display by Kristian Bell. Another beautiful morning on a beautiful beach on the Gulf Coast of Florida seemed to prompt these two royal terns to commence an intricate courtship dance. The photograph was taken with a Canon 300mm lens and 2x extender. (Photo by Kristian Bell/Deakin University/Royal Society Publishing Photography Competition 2018)

Runner-up. “The heavens opened and the streets around Shibuya, Tokyo, were suddenly drenched in even more colour and movement. I was taking photos (my friend was holding the umbrella) when I noticed this woman at the crossing. I’m pleased with the futuristic feel of the image”. (Photo by Katherine Bridgestock/The Guardian)

Elena Chernyshova's vision of Norilsk, Russia, the northernmost city in the world, is a series of surprises by which she extracts otherworldly beauty from ugly realities. Here: 2013. A woman is visible through a narrow passageway between two buildings. Norilsk's urban spaces were designed to shorten distances around large developments and give residents maximum protection from arctic winds. (Photo by Elena Chernyshova)

A young Shiite Muslim child participates in a Muharram procession in Ahmadabad, India, Sunday, October 9, 2016. (Photo by Ajit Solanki/AP Photo)

A cooling tower is seen under mechanical demolition in Binzhou, Shandong Province, China, July 13, 2016. (Photo by Reuters/China Daily)

Korean War veteran Jon Sun Rye, 83 years old, walks after she paid respect to former leaders Kim Il-sung and Kim Jong-il at Mansu Hill in Pyongyang, North Korea, 15 April 2016. North Korea is marking the “Day of the Sun”, celebrating the day of birth of the country's founder, Kim Il-sung. The Day of the Sun is a national holiday. (Photo by Franck Robichon/EPA)

Angel Paz, dressed as a calavera, waits with family for the start of the Gran Procession of the Catrinas, to mark the upcoming Day of the Dead holiday, in Mexico City, Sunday, October 23 2016. (Photo by Anita Baca/AP Photo)