These portraits reveal the incredibly humanlike expressions of a variety of apes.Through piercing eyes and finite facial details, the intimate photographs show the animals looking angry, sad, delighted and pensive. They are the works of Manuela Kulpa – an IT consultant and keen photographer from near Cologne, Germany – who shot the apes predominantly at zoos across Austria, Belgium, the Czech Republic, Germany and the Netherlands. Here: Bonobo, Azibo. (Photo by Manuela Kulpa/Caters News)

Runner-up. “The heavens opened and the streets around Shibuya, Tokyo, were suddenly drenched in even more colour and movement. I was taking photos (my friend was holding the umbrella) when I noticed this woman at the crossing. I’m pleased with the futuristic feel of the image”. (Photo by Katherine Bridgestock/The Guardian)

A ring-tailed lemur is pictured at Zoom Torino, a zoological park in Cumiana near Turin, on April 22, 2015. Zoom Torino is a new immersive zoological park, where animals can be seen without bars or cages, only natural barriers ensure the visit. (Photo by Marco Bertorello/AFP Photo)

These powerful images capture the spear-wielding tribes of Papua new Guinea who believe they are possessed with the spirit of the crocodile. They show how the Kangunaman clansmen scar their backs to resemble reptile scales while the Huli Wigmen wear elaborate headdresses to signal they are ready for battle. (Photo by Trevor Cole/Media Drum World)

Youth category winner: Symphony No 5 by Giacomo Redaelli (Italy). (Photo by Giacomo Redaelli/2019 Nature Photographer of the Year)

“This grey adult male seal enjoys a good grin while lying on the beach”. (Photo by Michael Hutch/SplashdownDirect via Rex USA)

German flying ace Baron Manfred von Richthofen (1892–1918) also known as The Red Baron with his dog Moritz, circa 1916.