Winner – Animal Portraits: The pose by Mogens Trolle, Denmark. A young male proboscis monkey cocks his head slightly and closes his eyes. Unexpected pale blue eyelids now complement his immaculately groomed auburn hair. He poses for a few seconds as if in meditation. He is a wild visitor to the feeding station at Labuk Bay Proboscis Monkey Sanctuary in Sabah, Borneo – “the most laid-back character”, says Trolle, “quite unlike anything I’ve ever seen on another monkey” – connects us, he hopes, with a fellow primate. (Photo by Mogens Trolle/Wildlife Photographer of the Year 2020)

A refugee sits in front of a Greek riot police cordon during a protest at the Greek-Macedonian border, near the Greek village of Idomeni, March 1, 2016. (Photo by Alexandros Avramidis/Reuters)

Winner, photojournalism. Elephant in the room, by Adam Oswell, Australia Zoo. Visitors watch a young elephant performing underwater. Oswell was disturbed by this scene, and organisations concerned with the welfare of captive elephants say performances like this encourage unnatural behaviour. In Thailand, there are now more elephants in captivity than in the wild. With the Covid pandemic causing tourism to collapse, elephant sanctuaries are becoming overwhelmed with animals that can no longer be looked after by their owners. (Photo by Adam Oswell/Wildlife Photographer of the Year 2021)

Wide-angle category winner. Part of the Illusion by Marcus Blatchford (UK). Location: National Dive & Activity Centre, Chepstow, Wales. “This is the deepest inland dive centre in the UK. The dive plan was to explore the deep end, but this time I dived ‘unplugged’ (without my strobes). With the exception of this change to my camera technique, there were no planned shots I wanted to achieve – just a fun dive with ad-hoc photos along the way. Shortly after this photo was captured, in 6C water and two hours of decompression ahead of us, we turned and started the long ascent back to the surface”. (Photo by Marcus Blatchford/Underwater Photographer of the Year 2016)

This perfectly-timed GoPro snap captures the moment an angrymagpiecame within inches of a biker, as swooping season takes hold in Australia. The notoriously aggressive bird can be seen flying with its wings tucked in so it looks exactly like a torpedo in the magnificent photograph from Middlemount, Queensland. Retail businesswoman Monique Newton, 53 was riding pillion with a friend when she spied the divebombing bird – but rather than speed off, they slowed down to capture it up close. (Photo by Monique Newton/Caters News Agency)

A model is seen posing in front of Olivier Rousteing, during the Balmain Pre-Fall 2021 Collection photoshoot by Olivier Rousteing, from Saint-Honore Flagship store in Paris, on December 09, 2020 in Paris, France. (Photo by Edward Berthelot/Getty Images)

Giant panda Shuixiu's two babies are pictured at their room in Shenshuping base of China Conservation and Research Center for the Giant Panda on November 3, 2021 in Aba Tibetan and Qiang Autonomous Prefecture, Sichuan Province of China. (Photo by An Yuan/China News Service via Getty Images)

A brown bears cools off in a pool at the bear sanctuary near the village of Mramor, on August 18, 2020. The bear sanctuary was closed for visitors on August 18, as part of safety measures to curb the spread of the COVID-19 (novel coronavirus) disease. (Photo by Armend Nimani/AFP Photo)