“When We're Gone... Memories Of Us Are All That Remain”, 2011. (Photo by: Trini61; Source: Flickr)

“Sperm whale opening it's huge mouth, Ogasawara Islands, Japan. Sperm whales are the biggest carnivorous animals on the planet. Each teeth in the mouth of adult specimen weights more than 1 kilogram. Interestingly enough that modern marine biologists believe that these teeth despite being fearsome play little role in capturing and eating giant squid – with their main function being mainly ritual aggression between males!”. (Photo by Alexander Safonov)

Jack London was a prolific photographer in addition to his writing. Here: Salvation Army barracks in London during Sunday morning rush – men who had been given tickets during the night queuing for free breakfast, 1902. (Photo by Jack London/Courtesy of Contrasto)

Runner-up. “The heavens opened and the streets around Shibuya, Tokyo, were suddenly drenched in even more colour and movement. I was taking photos (my friend was holding the umbrella) when I noticed this woman at the crossing. I’m pleased with the futuristic feel of the image”. (Photo by Katherine Bridgestock/The Guardian)

In this handout image provided by the City of New York, a Nissan NV200 is seen as a New York City taxi cab in New York City. The Taxi and Limousine Commission along with New York City Mayor Michael Bloomberg announced the Nissan NV200 designed by Nissan North America, Inc., has been chosen as the winner of the Taxi of Tomorrow competition and will become the City's exclusive taxicab for a decade. (Photo by City of New York via Getty Images)

Honourable mention, Behaviour category. Toss the scorpion – Indian roller playing with its kill by Susmita Datta. The image was taken during an early morning safari drive at Tadoba Andhari Tiger Reserve in India. (Photo by Susmita Datta/PA Wire/Royal Society Publishing Photography Competition 2017)

Winner. “I took this vertical image in the Quarry Bay district of Hong Kong during the dusk ‘blue hour’, when there was a perfect balance between the ambient light in the sky and the artificial lights of the high-rise residential buildings”. (Photo by Jatinder Heer/The Guardian)

Behaviour winner; Courting Royals: two royal terns in courtship display by Kristian Bell. Another beautiful morning on a beautiful beach on the Gulf Coast of Florida seemed to prompt these two royal terns to commence an intricate courtship dance. The photograph was taken with a Canon 300mm lens and 2x extender. (Photo by Kristian Bell/Deakin University/Royal Society Publishing Photography Competition 2018)